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The Servamps and their pact-bound "Eves" are finally getting back to their normal lives as they recover from their injuries from the previous battles. However, when it starts snowing in the middle of summer, one of the Eves, Mahiru Shirota, suspects vampiric interference. Concerned by the strange phenomenon, he sets out to gather the group once more to try and solve the mystery; however, they suddenly lose contact with Misono Arisuin, the Eve of the Servamp of Lust. Servamp Movie: Alice in the Garden delves into the untold past of Misono and his brother Mikuni Arisuin, as well as the many mysteries of the grand Arisuin Mansion.

The story revolves around a 16-year-old boy (15 at the beginning of the story) named Mahiru Shirota who likes simple things and dislikes difficult things. One day, he picks up a stray black cat he finds in the middle of the road and gives it the name Kuro (which literally translates into "black"). From this moment on, a contract between the servamp Sleepy Ash and Mahiru is formed, and Mahiru is pulled into the Servamp war between the seven deadly Servamps and their eighth sibling Tsubaki.